- Ogura et al. (2003) .A Case of Accidental Death of a Kerama Deer (Cervus nippon keramae) in Tokashiki Island. Jpn J Zoo Wildlife Med. 8(1), 55-62 (in Jpn with English abst)
- Ohtaishi (1986) Preliminary memorandum of classification, distribution, and geographic variation on Sika deer. Mammal Sci. 53, 13-17 (in Jpn)
- Tamate et al. (2000) .Genetic Differentiation among Subspecies of the Sika Deer (Cervus nippon), with Special Reference to the phylogeny of C. n. keramae in the Kerama Island Group. Tropics. 10(1), 73-78.
- etc.
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This subspecies may originate from an artifical introduction, and the small size may be an adaptation to the smal island environment. Paleographic studies suggested the age of introduction should be early 17 C. A genetic study using mitochondrial DNA sequences clarified that there is little genetic difference between this subspecies and Kyushu population of Cervus nippon.
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