Rana rugosa

Basic information
Scientific name	Rana rugosa	Click to magnify
Major synonym	Glandirana rugosa
Common names	wrinkled frog
Higher taxon	Ranidae, Anura, Amphibia
Natural range	Northeastern continental China, Korean Peninsula, coastal area of far eastern Russia, Japan. In Japan, Honshu, Shikoku, Kyushu, and peripheral islands (Sadogashima, Oki, Iki, Goto Islands, Yakushima, Tanegashima, etc.)
Habitat	Rice field, artifical pond, lowland and montane stream, swamp, etc.

Invasion information
Range in Japan	Izu Islnads (Izuoshima, Niijima, Miyakejima), Hokkaido (several areas in southern and western part). There is a record in Kuchinoshima Is. (Tokara Islands) where establishment status is unknown.
Origin	Unknown. Possibly several areas in Japan.
Date	The first record in Hokkaido was in 1985.
Route	Accidental: Tadpole of this species may have hitchhiked on Carp seedling for aquaculture (Hokkaido population).
Impact	Unknown Native organism(s) affected:
Regulation in Japan	No action for prevention, mitigation, control, or eradication.
Introduced range in other countries	Hawaii (from Japan)

Reference

Notes

Frost et al. (1998-2010) Amphibian Species of the World (Accessed on 2012-6-25) http://research.amnh.org/herpetology/amphibia/.
Hasegawa (1999) Impacts of the introduced weasel on the insular food webs. pp.129-154. Ota (編) Tropical Island Herpetofauna: Origin, Current Diversity, and Conservation. Elsevier Science B. V., Amsterdam.
Maeda & Matsui (1999) Frogs and Toads of Japan. Bun-ichi Sogo Shuppan, Tokyo (in Jpn)
Maenosono & Toda (2007) Distribution of amphibians and terrestrial reptiles in the Ryukyu Archipelago: A review of published records. Akamata. 18, 28-46 (in Jpn)
Matsuo (2004) Amphibians and Reptiles of Nagasaki Prefecture, Japan. Nagasaki Shinbunsha (in Jpn)
Sengoku et al. (eds) (1996) The Encyclopedia of Animals in Japan 5 Amphibians, Reptiles, Chondrichthyes. Heibonsha, Tokyo (in Jpn)
Tokuda (2011) Compact Picture Guide to Hokkaido Reptiles & Amphibians. Hokkaidoshinbunsha, Sapporo (in Jpn)